The figure was revealed by UKAD following a Freedom of Information (FOI) request made to the National Anti-Doping Organisation.

The majority of the figure was made up of legal costs, with £576,587 ($800,000/€648,000) in fees going to legal firm Bird & Bird LLP, while a further £1,130 ($1,500/€1,200) was paid to a barrister.

"Bird & Bird LLP was instructed by UKAD throughout the Fury cases and so the fees paid to that firm were in respect of advice on an ongoing basis, all aspects of case preparation and representing UKAD in front of the National Anti-Doping Panel tribunal," UKAD stated.

"The barrister provided one piece of written advice."

Laboratory analysis and associated services also cost £7,942 ($11,000/€8,900), giving the cases a combined cost of £585,659 ($813,000/€658,000).

UKAD said they are looking to recover £250,000 ($347,000/€281,000) from their insurer, with the net cost therefore expected to be £335,659 ($466,000/€377,000).

The organisation has a budget of around £8 million ($11.1 million/€9 million), which comes from both Government funding and commercial income.

There had been concerns raised in November that should legal action be taken by Tyson Fury, UKAD could be made bankrupt.

Following the FOI request, it was claimed there "was never any danger that UKAD would become insolvent".

The cases ultimately led to both fighters accepting backdated two year bans in December.

It meant Tyson and his cousin Hughie were free to fight again if they received a renewed licence from the British Boxing Board of Control.

"The money spent by UK Anti-Doping in the Fury case shows that if we determine there is evidence of doping, we will pursue a case against an athlete, coach or doctor, regardless of their public profile or status," said Nicole Sapstead, UKAD chief executive.

"In this case, two Anti-Doping Rule Violations were upheld and two-year bans given to each athlete.

"As an arm's length body of Government, we are always careful about how we spend public money, and the consistent support of the UKAD Board and the Department of Digital, Culture, Media and Sport ensures our legal decisions are made for the right reasons and not financial ones."

Tyson Fury had been charged by UKAD after testing positive for a "prohibited substance" - the anabolic steroid nandrolone - in June 2016 but he claimed that was a result of eating a wild boar.

The former world heavyweight champion failed to attend the hearing in December.

UKAD, Tyson and Hughie Fury then released a joint statement which stated all parties had agreed to resolve proceedings and bring the case to an end.

Twenty-nine-year-old Tyson Fury, from Manchester, shocked the boxing world in November 2015 when he defeated Wladimir Klitschko by a unanimous points decision after 12 rounds in Düsseldorf, to win four world heavyweight belt.

A rematch with Klitschko was scheduled for July 2016 in Manchester but was subsequently postponed after Fury claimed to have suffered an ankle injury before details of the positive drugs test emerged.